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Princess Luna Approves Greatly.
Nerd.
:þ
In old english, nominative/accusative/genitive singular “brōþor” and dative singular “brēþer”.
brōþor -> brother
brēþer + -en -> brethren (common in middle english)
brōþor + -s -> brothers (rare in middle english)
(þ is pronounced as modern th)
Base word probably became “brother”. “Brother-en”.
“Brethren” is already plural, no need for an “S” at the end. I believe it’s an old plural form, similar to “children” and “oxen”, two of the few remaining “-en” plural forms in the English language.